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The mutation rate of an organism is highly variable. Some species of bacteria and viruses have mutation rates of up to 10^-8^ per base pair. Other microorganisms have mutation rates as low as 10^-10^. For example, the mutation rate for *Salmonella* is approximately 3×10^-10^ (Abbitt and Leffler, 1992), *Escherichia coli* is approximately 5×10^-10^ (Lea and McCarty, 1990), and yeast cells are approximately 5×10^-11^ per base pair (Cairns, 1974). Within the cell, the rate of mutation is dependent on many factors, such as the rate of transcription, the rate of 0b46394aab